I just happened to see this article today and wanted to share. Dr. Al has devoted his life to breeding hops and this may help answer many of the questions folks have about hops in general along with some solid breeding information.
http://faculty.agron.iastate.edu/fehr/HOCP/27HOCP.pdf
http://faculty.agron.iastate.edu/fehr/HOCP/27HOCP.pdf