Underletting is the best way to exclude the most air from being trapped in your grist when adding the water. Slowly filling from the bottom, the water uniformly displaces the air with minimal bubbles adhering to the crushed grain kernels. However if your crush is too fine then the flour and small particles will hinder the uniform wetting of the grain creating dry pockets and dough balls. With a stirred hido mash this is of little concern but when running a low oxygen recirculating system, trapped air and doughballs are detrimental. In any recirculated system too fine a crush is bad as it will cause stuck mashes, channeling, poor flow, compressed grain bed, low efficiency etc. To get the best grind for recirculated mash, not only do you want a coarser gap but also be milling at a slower RPM. Below 100 is good but below 70 is optimal.
Lastly, for best results when underletting, you want to keep your strike water temperature below the gelatinization temperature of your grain. I usually shoot for an initial water temperature 140 or below, then after filling proceed to your first rest temperature.