I do think the acetic acid mostly comes from brett. I came to this realization on my own (if anyone but me take my realizations for anything), when I read about a brewery souring by oxygenating brett only beers in barrels. I cannot remember the brewery name but in the presence of oxygen brett will produce acetic acid. I have not found a metabolic pathway map for brett (I haven't look real hard though), if I could find one it would explain everything. If one does not exist who knows then. A metabolic pathway map would demonstrate what is made in the presence of oxygen versus anaerobic conditions. I'm trying to figure these things out and do research but even with my university library has limited offering on Brettanomyces behavior. A large majority of academic research papers is directly related to the wine industry with the only beer references from 20yrs ago from Leuven about lambics.
When I find anything cool I'll post up and share the info.
When I find anything cool I'll post up and share the info.