Keep it cold, and not too acidic.
Comprehensive Sensomics Analysis of Hop-Derived
Compounds during Storage of Beer
Daniel Intelmann, Gesa Haseleu, Andreas Dunkel, Annika Lagemann, Andreas Stephan and
Journal of Agricultural and Food Chemistry,59, 1939-1953 (2011)
The authors tracked the concentration of nearly 70 hop constituents
and their isomerization products in packaged beer over 8 months,
4 years and 10 years, and at different storage temperatures and
pH's. Their conclusion:
"The sensomics data obtained during this study
offers the scientific basis for a knowledge-based optimization of
the beer bitter taste by controlling the initial pH value of the beer
by technological means and by keeping the temperature as low as
possible during storage of the final beverage."
By pH value they mean a small difference makes a large difference
in the amount of harsh bitter compounds that form in the beer
(pH 4.4 is much better than pH 4.2).
Ask not for whom the beer twangs; it twangs for thee.